The Genesect are Pokémon resurrected by the Plasma Gang through the modifiation of 300 million year old fossils. When they return to the place they once knew as their home and find it has been radically changed, now being a mountain ravaged by blizzards, they lose themselves in despair.
"Want to... go home..." With these words, the five Genesect head towards the metropolis New Tork City, searching for the 300 million year old landscape they once knew.
Mewtwo sensed the Genesect's voices. Just like the Genesect, Mewtwo is a Pokémon created by human hand; a Pokémon born from the genes of Mew.
At that point, Satoshi, Pikachu and their travelmates are visiting New Tork City. They have come there to check out "Pokémon Hills", a facility created to protect and shelter the Pokémon in the area. They are amazed at how Pokémon can live peacefully in the middle of a giant city like this.
Then, the Genesect arrive. The red Genesect fires an energy blast towards Satoshi and the other with incredible speed! "Eliminate the enemy!", it says.
Incredibly, the group is saved from certain death by the legendary Pokémon Mewtwo. "Genesect are Pokémon not meant to exist in this world. They're just like me", it says. Understanding perfectly well their grief, Mewtwo confronts the Genesect as they attack the city. Fierce shockwaves from their intense brawl fly through the entire city.
As the two Pokémon both surpass their limits, the battle turns into the wildest Pokémon speed battle ever seen! What is the true power displayed by the Awakened Mewtwo? And how will this destined battle end?
Satoshi has vowed to take the Genesect to their home. Alongside Pikachu, he sets off to help the city through this crisis. Will he manage to fulfill his promise!?
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For maximum enjoyment, we recommend that you watch the Highlight Showcase Special and the tie-in TV special before the movie.